1981-86 : National Veterinary School of Maisons Alfort (France) (Diploma obtained in September 1986).
1986-87 : DEA (Master degree of Science ) of Molecular and Cellular Biology of Development : University of Paris 6 (Diploma obtained in October 1987)
1987-92 : Ph.D., University of Paris 6 (Thesis defended in June 1992)
2000 : Course of Mouse Genetic (Pasteur Institute)
2011 : Habilitation à Diriger les Recherches (official authorisation to supervise PhD. students)